TYPE: Costa Rica. [San José. Carrillo,] La Palma, 1500 m, August 1910, C. Wercklé s.n. [840 in Herb. O. Jiménez] (holotype, B, destroyed; lectotype, designated by Pupulin (2010), drawing of the holotype made under the supervision of Schlechter, AMES; photograph of the holotype ex B, AMES).