TYPE: San José: Pérez Zeledón, San Ramón Norte, trail to Cerro Pelón, 9°25'N 83°44'W, 1050 m, lower montane moist forest, secondary mature and primary vegetation, collected by J. Cambronero and F. Pupulin, flowered in cultivation, 3 December 2001, F. Pupulin 3499 (holotype, USJ; illustration of type).