TYPE: Ecuador. Pastaza: north of Topo, epiphytic in forest along Río Zuñag, alt. 2200 m, 23 Feb 1990. C. Luer illustr. 14722, A. Hirtz, S. Dalström, L. Årnby, J. Del Hierro, E. Astorga, S. Ortega & J. Cabezas 4647 (holotype, MO; illustration of type).