TYPE: Costa Rica. Boundary between Alajuela and Heredia provinces: Grecia, Sarapiquí, Colonia Virgen del Socorro, road to Cariblanco, bridge on San Fernando river, 10º16'32"N, 84º10'16"W, 750 m, shores of San Fernando river, 13 Feb 2004, D. Bogarín, H. León-Páez, F. Pupulin & E. Salas 741 (holotype, JBL-spirit; isotypes, CR, JBL-spirit; illustration of type).