TYPE: Ecuador. Pastaza: north of Topo, epiphytic in forest along Río Zuñig, alt. 2200 m, 23 February 1990, A. Hirtz, S. Dalström, L. Årby, J. Del Hierro, E. Astorga, S. Ortega & J. Cabezas 4662 (holotype and illustration of type, MO; illustration of type).