SYNTYPES: Ecuador. Pichincha. In silvis subandinis prope Canzacoto, Sodiro 146 (B, destroyed); Juli 1883, ad terram, in silvis interandinis, Mille 32 (B, destroyed). Ecuador: Zamora-Chinchipe: Yangana to Valladolid, beyond the pass, 1950 m, 24 July 1985, C.H. Dodson 16054, lectotype [neotype?] designated by Dodson (1994).