TYPE: Nicaragua. Granada: falda W Volcán Mombacho, road and trail above Finca Santa Ana from reservoir to somewhat above Plan del Flores, ca. 950-1150 m, 11.49-50N 85.58W, disturbed cloud forest below, elfin forest above, 1 Oct 1977, W.D. Stevens 4331 & B.A. Krukoff (holotype, MO; isotype, CR).